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Review Article Volume 15 Issue 1
Paleontology, stratigraphy and paleogeography of ninety African Campanian-Neogene planktic and benthic foraminifera of Anan
Haidar Salim Anan
Received: December 31, 2025 | Published: March 2, 2026
Citation: Anan HS. Paleontology, stratigraphy and paleogeography of ninety African Campanian-Neogene planktic and benthic foraminifera of Anan. J Aquac Mar Biol. 2026;15(1):27-36. DOI: 10.15406/jamb.2026.15.00438
Abstract
This study deals with taxonomic consideration of ninety African Campanian- Neogene benthic and planktic foraminiferal species of Anan, which belongs to fifty-five genera. Rich and well-preserved African fauna from Morocco, Tunisia, Egypt, Ivorian Basin, Nigeria and Tanzania, made it possible to correlate them with those previously identified species in different stratigraphic sequence in many Tethyan localities: USA, Trinidad, Spain, France, Germany, Hungaria, Jordan, Iraq, United Arab Emirates (UAE), Qatar and India. Four new benthic foraminiferal genera were erected by the present author: Leroyia1 (3 species), Lenticuzonaria2 (1 species), Lenticubella3 (2 species) and Percultalina4 (2 species). 90 benthic foraminiferal species: 15 agglutinated species, 2 porcelaneous, 34 Lagenid, 33 Rotaliid, and 6 planktic species are added to this assemblage. 70 species of them (~78%) are recorded from Egypt, 9 species (~10%) from Tunisia, 6 species (~7%) from Ivorian Basin, 5 2 species (~2%) from each of Nigeria and Tanzania, and only one species (~1.5%) from Morocco. One of the recorded species is believed here to be new: Oolina tunisica. This study presented also five benthic foraminiferal evolutionary lineages: Textulariel la cretosa Cushman > T. sinaensis Anan, Leroyia maqfiensis Anan > L. aegyptiaca Anan, Percultalina misrensis Anan > P . sinaensis Anan, Uvigerinella tunisica Anan > U. nakkadyi Anan, Anomalinoides aegyptiacus (LeRoy) > A. dababiyaensis Anan, and another four planktic lineages : Plummerita safaae Anan > P . haggagae Anan, P. salimi > P . hodae, Anan, P . inflata (Brönnimann) > P . costata (Brönnimann), and P. tunisica Anan > P . spainica Anan lineages. The Campanian Gaudryina jaffi Anan5 and Early Eocene G. ameeri Anan,6 Campanian Gaudryina lawai Anan7 and G. speijeri Anan,6 Campanian Pseudoclavulina farisi Anan,7 and Paleocene P . hewaidyi Anan, 8,9 Middle Eocene Hemirobulina bassiounii Anan 10 and Oligocene-Miocene H. hantkeni (Bandy) have good examples of Homeomorphy.
Keywords: Paleontology, foraminifera, stratigraphy, lineage, homeomorphy, campanian, paleogene, neogene, Africa
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